Sambucus racemosa, or red elderberry, shows potential for regenerative agriculture, though current knowledge base coverage is limited. Based on available information, its primary roles appear to be as a component in polyculture systems and as a potential forage source. Regenerative benefits may include support for pollinators and potentially contributing to soil health through its biomass. While specific integration with practices like rotational grazing or no-till is not detailed in the knowledge base, its shrubby nature suggests it could fit within agroforestry designs or as a hedgerow species. Direct farmer experiences or insights into its effectiveness as a nitrogen fixer, cover crop, or carbon sequesterer within regenerative systems are not extensively documented in the provided material. Further research and on-farm observation are needed to fully understand its contributions to soil building and broader ecosystem services in regenerative contexts.

Regenerative Quick Profile

All recommendations assume integrated, regenerative practices—not conventional inputs.

Climate & Soil Fit

Climate: Tropical Rainforest, Tropical Monsoon, Tropical Savanna, Hot Semi-Arid (Steppe), Cold Semi-Arid (Steppe), Hot Desert, Cold Desert, Humid Subtropical, Oceanic (Maritime Temperate), Hot-Summer Mediterranean, Warm-Summer Mediterranean, Monsoon-Influenced Humid Subtropical, Subtropical Highland, Hot-Summer Continental, Warm-Summer Continental, Subarctic, Monsoon-Influenced Hot-Summer Continental, Tundra

Zones: USDA 3-7, Australian Zones 4-6

Optimal Soil: Loam Soil

System Role & Functions

Primary: Pollinator Support

Secondary: Forage Integration, Cover Crop System

Key Benefits: Fast production, Integration-friendly

Management Level

Experience: Beginner-Friendly

Maintenance: Moderate maintenance - Red elderberry is adaptable and low maintenance, with pruning integrated for plant health and occasional compost application to support ongoing fertility and fruiting.

Time to Production: Fast (1-2 years) - This elderberry provides early harvests within 1-2 years, with good fruit yield by year 2-3, contributing to a resilient food system.

Value Streams

  • Fruit/nut harvest
  • Livestock forage value
1

Climate Suitability Assessment

Will this plant thrive in your climate?

IDEALLY SUITED

Köppen Zone: Cfa (Humid Subtropical), Cfb (Oceanic (Maritime Temperate)), Dfa (Hot-Summer Continental), Dfb (Warm-Summer Continental)
USDA Zone: 5a, 5b, 6a, 7a, 8a, 9a
Australian Zone: temperate
EU Climate Region: atlantic

Red Elderberry thrives in climates with mild winters and long, warm growing seasons, typically experiencing fewer than 10-15°F (-12 to -9°C) winter lows and ample rainfall (30-50 inches/75-125 cm annually). These conditions are met in USDA Zones 5b-8b, Australian temperate zones, and the EU Atlantic climate region. The plant establishes easily, flowers profusely to support pollinators, and produces abundant fruit for forage integration. Its growth cycle aligns perfectly with these temperate to warm-temperate climates, requiring minimal management beyond basic pruning. The species exhibits excellent vigor and resilience, contributing significantly to biodiversity and ecosystem services within regenerative agriculture systems. Its ability to adapt to various soil types, provided they are well-drained, further enhances its suitability in these regions, ensuring reliable performance year after year.

ADEQUATE

Köppen Zone: BSk (Cold Semi-Arid (Steppe)), Csa (Hot-Summer Mediterranean), Csb (Warm-Summer Mediterranean), Cwa (Monsoon-Influenced Humid Subtropical), Cwb (Subtropical Highland), Dfc (Subarctic), Dwa (Monsoon-Influenced Hot-Summer Continental)
USDA Zone: 4a, 10a, 11a

Red Elderberry can perform adequately in climates with cold winters (down to 0°F/-18°C) and moderate growing seasons, such as USDA Zones 4b-5a and 9a-9b, and some parts of the EU Atlantic region. In colder zones, winter survival is generally good, but fruit production might be reduced by early frosts or a shorter ripening period. In warmer zones (9a-9b), summer heat and potential drought can stress the plant, necessitating supplemental irrigation and possibly afternoon shade to maintain vigor and fruit yield. While not performing at its peak, it still offers valuable pollinator support and forage integration. Management may involve protecting young plants from extreme cold or ensuring adequate water during hot spells, making it a viable, though not optimal, choice for regenerative agriculture in these transitional climates.

NOT RECOMMENDED

Köppen Zone: Af (Tropical Rainforest), Am (Tropical Monsoon), Aw (Tropical Savanna), ET (Tundra), BSh (Hot Semi-Arid (Steppe)), BWh (Hot Desert), BWk (Cold Desert)
USDA Zone: 2a, 3a, 3b, 12a

Red Elderberry is not recommended for climates with extreme winter cold (below 0°F/-18°C) and very short growing seasons, or for hot, arid regions. This includes USDA Zones 1a-4a, Köppen Dfc, Dwc, and Dwd. In very cold zones, winter kill is highly probable, and the limited frost-free period prevents adequate growth and fruit maturation. Establishment success is very low, and consistent yields are impossible. In hot, dry climates, the plant struggles with heat stress, requiring extensive irrigation and suffering reduced vigor and fruit production. The combination of extreme temperatures (both cold and hot), insufficient growing season length, and water stress makes Red Elderberry economically and practically unviable in these zones. Alternative, more resilient species adapted to these specific harsh conditions are essential for successful regenerative agriculture.

Better alternatives for these "not recommended" zones: Saskatoon Berry (Amelanchier alnifolia) (Highly cold-hardy and adapted to continental extremes, native to many cold regions.), Highbush Cranberry (Viburnum trilobum) (Extremely cold-hardy shrub with edible berries, tolerates a wide range of conditions.), Lingonberry (Vaccinium vitis-idaea) (Extremely cold-hardy, suitable for short growing seasons and acidic soils.), Buffaloberry (Shepherdia argentea) (Drought-tolerant native shrub with edible berries, thrives in cold, arid environments.)

Note: Zones listed above represent climates where this plant can produce reliably with reasonable management. Climate zones not mentioned would require intensive climate modification (greenhouses, extensive infrastructure) and are not economically viable for regenerative agriculture purposes.

2

Soil Suitability Assessment

Which soil types work best for this plant?

IDEALLY SUITED

Loam Soil

This plant thrives in these soil types without requiring amendments or remediation. Natural soil conditions support optimal growth and productivity.

ADEQUATE

Clay Soil, Rich Soil, Rocky Soil, Sandy Soil

This plant performs acceptably in these soil types with moderate, manageable remediation such as pH adjustment, compost addition, or drainage improvement. The required amendments are practical and cost-effective for regenerative agriculture.

NOT RECOMMENDED

Acidic Soil, Alkaline Soil, Desert Soil, Saline Soil, Wet Soil

Growing this plant in these soil types would require impractical remediation such as complete soil replacement, extensive amendments, or cost-prohibitive infrastructure. These conditions are not economically viable for regenerative agriculture.

Note: Soil suitability assessments focus on remediation requirements. "Ideally Suited" means the plant generally thrives without the need for substantial amendments, "Adequate" means manageable remediation (lime, compost, mulch), and "Not Recommended" means impractical soil changes would be required. Climate factors like rainfall and temperature also influence success.

3

Seasonal Considerations

Planting timing, growth duration, and harvest windows

Establishing red elderberry is a multi-year commitment. For nursery stock, bare-root trees are best planted during winter dormancy, typically in late fall or very early spring before bud break. Container-grown plants offer more flexibility, allowing planting throughout the active growing season, though early spring or early fall are ideal to minimize transplant shock.

Expect a few years for your red elderberry to become truly established, usually two to three years before you see a meaningful first harvest. Full production, where yields are consistent and abundant, will take another two to three years, with trees remaining productive for decades.

Seasonal management focuses on optimizing growth and harvest. Pruning is best undertaken during the winter dormancy period, allowing the plant to focus energy on new growth in spring. Bloom typically occurs in late spring or early summer, followed by fruit development through summer. Harvest usually follows in mid to late summer, depending on your specific climate and variety. Throughout the year, observe your plants, noting their response to weather patterns and ensuring they enter a robust winter dormancy to prepare for the next productive cycle.

4

System Role & Multi-Benefit Value

Functional roles, integration strategies, and stacked benefits

Functional Role

Total System Value

Red elderberry offers substantial system value beyond direct harvest, primarily through its role in enhancing farm biodiversity and ecosystem services. Its early blooming flowers are a critical resource for pollinators at a time when other food sources may be scarce, directly supporting crop pollination and natural pest control. As a shrub, it contributes to soil health by stabilizing slopes and reducing erosion, particularly in less-managed areas like hedgerows or riparian buffers. Its berries provide food for birds and small mammals, further diversifying wildlife on the farm. While direct harvest value from the berries can be realized (though often secondary to wildlife consumption), the plant's greatest contribution lies in its support of a complex, resilient agroecosystem. This benefit stacking enhances overall farm health, reduces reliance on external inputs, and builds ecological stability.

Integration Characteristics

Multi-Benefit Value: Adequate - Offers berries for wildlife and human use, with moderate pollinator attraction, contributing to ecological diversity and farm resilience.

Integration Friendliness: Ideally Suited - Similar to S. nigra, this elderberry integrates well into diverse farm systems, providing berries and flowers while tolerating interplanting and supporting wildlife.

Sources behind this view

Videos & Podcasts
5

Management & Care Requirements

Integration guidance, maintenance needs, and care practices

How to Integrate This Plant

Red elderberry (Sambucus racemosa) is a versatile shrub that can be integrated into regenerative farm systems primarily for pollinator support and wildlife habitat. As a non-tree shrub, it excels in edge habitats, hedgerows, and as understory planting in silvopastures or food forests. Its early spring blooms provide crucial nectar and pollen for emerging insects, making it a valuable component in supporting beneficial insect populations. It can also contribute to erosion control on slopes and in riparian buffer zones. Compatible practices include food forests, silvopasture (as a non-browseable understory or edge planting), and hedgerows. Timeline to contribution is relatively quick, with significant pollinator support starting in Year 1 and fruit production becoming noticeable by Year 3-5. Multi-benefit stacking includes habitat for birds and small mammals, soil stabilization, and aesthetic value, all contributing to a more resilient and biodiverse farm ecosystem.

Integration Practices & Management

Information regarding the specific integration methods of Sambucus racemosa (Red Elderberry) within regenerative agriculture systems is limited within the provided knowledge base. The available sources do not detail establishment techniques such as seeding rates, optimal timing, companion planting strategies, or the use of no-till versus minimal tillage for this species. Similarly, direct insights into its integration with grazing practices, including mob or rotational grazing, specific timings, or necessary rest periods, are not present. Termination strategies, whether through natural winterkill, grazing down, crimping, mowing, or herbicide use, are also not elaborated upon. Management considerations like fertility requirements, competition management, or succession planning for Sambucus racemosa in regenerative contexts are not discussed. Furthermore, its integration with cash crops through relay cropping, intercropping, or rotation sequences is not described. Consequently, practical farmer experiences and specific insights from the knowledge base concerning the 'how' of integrating Sambucus racemosa are not available.

Management Profile

Maintenance Intensity: Adequate - Red elderberry is adaptable and low maintenance, with pruning integrated for plant health and occasional compost application to support ongoing fertility and fruiting.

Pest Disease Pressure: Adequate - Red elderberry demonstrates good resilience, generally remaining healthy within a balanced ecosystem that supports beneficial insects and natural pest regulation.

Time To Production: Ideally Suited - This elderberry provides early harvests within 1-2 years, with good fruit yield by year 2-3, contributing to a resilient food system.

Sources behind this view

Videos & Podcasts
6

Economics & Value Streams

Direct harvest, system benefits, ecosystem services, and risk diversification

Comprehensive economic analysis including direct harvest value, system enhancement contributions, ecosystem services, value timeline, and risk diversification strategies.

Per-Tree Production Economics

Metric Value
Establishment Cost $8-15
Years to First Harvest 2-3 years
Annual Maintenance $3-6
Yield 10-20 lbs/year 4-9 kg/year
Market Price $1-3/lb $3-6/kg
Productive Lifespan 10-15 years
Net Annual Return* $2-$56/year

Values shown per mature tree, not per acre. In regenerative systems, trees are integrated at low densities across diverse landscapes. Establishment costs spread over the lifespan of the tree. Early years have costs but no revenue.

* Net Annual Return = (Yield × Market Price) − (Amortized Establishment Cost + Annual Maintenance). This return is realized only at/after first harvest; early years have costs but no revenue. Range shows worst case to best case scenarios.

System Enhancement Value

Beyond harvest: pollination services for your crops and ecosystem

Pollination Service Provision

Red Elderberry (*Sambucus racemosa*) significantly enhances system value through its primary function of pollinator support, attracting a variety of beneficial insects essential for farm productivity. Its secondary function as forage integration means it can be incorporated into pasture systems, providing browse for livestock or wildlife without requiring dedicated monoculture. Furthermore, its role as a cover crop system, particularly when planted in hedgerows as suggested by for VELB management, contributes to soil health, erosion control, and habitat creation. While its berries are generally considered toxic for direct human consumption without careful processing, they offer a valuable food source for birds, contributing to wildlife habitat and natural pest control. The plant's ability to be propagated via live stakes or hardwood cuttings makes it an accessible and cost-effective option for establishing these multi-functional ecological services across the farm landscape, contributing to overall farm resilience and biodiversity.

Ecosystem Service Contributions

Environmental contributions: carbon, pollinators, wildlife, and water

  • Carbon Sequestration: As a shrub that can reach approximately 9ft in height, Red Elderberry contributes to carbon sequestration through biomass accumulation in its stems, leaves, and root system. Its perennial nature allows for sustained carbon storage over its lifespan, with potential for increased sequestration as plants mature and form dense stands.
  • Pollinator Support: High - Red Elderberry is explicitly noted for its value to pollinators, making it a crucial component for enhancing biodiversity and supporting the reproductive success of other crops and plants within the integrated farm system.
  • Wildlife Habitat: Provides valuable habitat and food sources for birds through its berries. Its dense growth habit can also offer nesting sites and shelter. While its berries require careful processing for human consumption due to toxicity, they are an important ecological resource for wildlife.
  • Water Quality: Not applicable

Value Timeline: Bloom & Establishment

When you'll see results: annuals bloom year 1, perennials mature 2-3 years

Years 1-2

Initial establishment of vegetative cover, contributing to erosion control. Early pollinator attraction as plants begin to flower. Potential for limited forage integration if browsed by wildlife or livestock.

Years 3-5

Established pollinator support with increased flowering and fruiting. Contribution to the cover crop system with improved soil structure and ground cover. Potential for increased forage integration as plants mature. Berry production for wildlife begins to become significant.

Years 10-20

Mature plant stature providing more substantial habitat and pollinator benefits. Dense growth may offer some windbreak or microclimate modification if planted in strategic locations. Full realization of its role in a cover crop or hedgerow system.

20+ Years

Long-term, stable ecosystem service provision including robust pollinator support, wildlife habitat, and soil health benefits. Continued contribution to the integrated farm system as a resilient perennial component.

Farm Risk Reduction

How pollinator support reduces crop failure risk

  • Multiple Revenue Streams: Pollinator support (indirectly increasing yields of other crops), wildlife habitat provision (potential for ecotourism or hunting leases), forage integration (livestock sustenance), cover crop benefits (soil health improvement, reduced erosion).
  • Temporal Income Spread: Ongoing ecosystem services (pollinator support, habitat) provided annually, with seasonal berry production for wildlife. Establishment of vegetative cover and soil improvement benefits accrue over time.
  • Market Risk Hedge: Reduces reliance on a single crop or market by providing multiple ecological and agronomic benefits that enhance the overall farm system's resilience. Its role in pollinator support can buffer against yield losses in insect-pollinated crops. Its integration into cover crop systems improves soil health, reducing the need for costly inputs and mitigating risks associated with soil degradation.

Sources behind this view

Videos & Podcasts
7

Regenerative Suitability Details

Comprehensive trait ratings for system integration assessment

Comparative ratings for this plant across key regenerative agriculture traits.

Trait Suitability Explanation
Drought Tolerance Adequate This elderberry variety exhibits moderate drought tolerance, thriving with mindful water management and ample moisture retention through mulching during dry periods.
Establishment Ease Adequate Red elderberry establishes readily within a regenerative system, benefiting from healthy soil biology and adapting well to a range of conditions.
Time To Production Ideally Suited This elderberry provides early harvests within 1-2 years, with good fruit yield by year 2-3, contributing to a resilient food system.
Multi Benefit Value Adequate Offers berries for wildlife and human use, with moderate pollinator attraction, contributing to ecological diversity and farm resilience.
Climate Adaptability Adequate Hardier than S. nigra, extending to zone 2, this elderberry prefers cooler, moist conditions, performing best where summer heat is moderated by the landscape.
Hardiness Zone Range Adequate More cold-hardy than S. nigra, thriving in zones 3-7, it offers robust cold tolerance but may benefit from microclimate considerations in areas with extreme summer heat.
Maintenance Intensity Adequate Red elderberry is adaptable and low maintenance, with pruning integrated for plant health and occasional compost application to support ongoing fertility and fruiting.
Pest Disease Pressure Adequate Red elderberry demonstrates good resilience, generally remaining healthy within a balanced ecosystem that supports beneficial insects and natural pest regulation.
Integration Friendliness Ideally Suited Similar to S. nigra, this elderberry integrates well into diverse farm systems, providing berries and flowers while tolerating interplanting and supporting wildlife.

Comparative System: Ratings compare plants within their economic category (e.g., cover crop nitrogen fixation compared to other cover crops, not to all plants). Individual farm conditions and management practices significantly influence actual performance.

8

Learn More

Why farmers use this plant and additional resources

Why Regenerative Farmers Use This Plant

Sambucus racemosa, commonly known as red elderberry, is a valuable perennial shrub or small tree that offers significant regenerative benefits within diverse agricultural systems. At maturity, it can sequester an estimated 2-5 tons CO2e/acre/year, contributing to long-term carbon drawdown through biomass accumulation and soil organic matter enhancement. Its extensive root system, which can reach depths of 6-15+ feet (1.8-4.5+ m), is highly effective at improving soil structure, enhancing water infiltration, and preventing erosion, particularly on slopes. Red elderberry begins to offer ecological services immediately upon establishment and can reach full canopy development within 5-10 years, providing consistent habitat and resource provision for beneficial wildlife and pollinators throughout its multi-decade lifespan. Its asset value accumulates over time as the plant matures, providing ongoing ecological and potentially economic returns.

Integrating Sambucus racemosa into agroforestry systems offers a suite of ecosystem services that enhance farm resilience. As a component of windbreaks or hedgerows, it effectively reduces wind speed, protecting crops and livestock, and moderating microclimates. Its dense foliage provides crucial habitat and food sources for a variety of birds and beneficial insects, supporting natural pest control. In silvopasture systems, its shade can offer relief to livestock during hot periods, and its presence can encourage biodiversity within the grazing area. The plant's ability to thrive in marginal or disturbed areas makes it an excellent choice for ecological restoration projects or for stabilizing buffer zones along waterways. It also serves as an excellent nurse crop or component in multi-story cropping systems, supporting biodiversity by providing habitat and food sources for beneficial insects and pollinators. Its vigorous growth habit can help suppress weeds, reducing the need for mechanical or chemical interventions. In silvopasture designs, the elderberry can offer shade and browse for livestock during warmer months, while its fallen leaves contribute organic matter to the soil. Companion planting with nitrogen-fixing species beneath its canopy can further enrich the soil, creating a symbiotic relationship that benefits the entire ecosystem.

The quantitative ecosystem benefits of Sambucus racemosa are substantial. Its flowers are a vital early-season nectar and pollen source for numerous pollinator species, including bees and butterflies, often supporting hundreds of visits per bloom during its flowering period. The berries provide critical food for migratory birds and other wildlife, particularly during late summer and autumn. The continuous addition of leaf litter and woody debris from mature plants significantly contributes to soil organic matter accumulation, estimated at 0.5-1.5 tons/acre/year of dry matter, which in turn improves soil water-holding capacity and nutrient cycling. This robust biomass production and decomposition cycle fosters a healthier, more active soil microbiome. Measurable soil carbon increases are often observed by year 5-7 as the plant matures and contributes significant biomass.

While not typically a primary cash crop for its berries in large-scale commercial operations due to processing challenges, the berries are edible and can be used for jams, jellies, wines, and medicinal tinctures, offering niche market opportunities and supplementary income. The plant's rapid growth and adaptability mean it can begin providing these benefits within a few years of establishment, with full production and ecosystem services realized within 3-7 years. Once established, it requires minimal annual inputs and continues to accrue ecological benefits and potential economic returns for 20-30 years or more.

Regional successes highlight the adaptability of Sambucus racemosa. In the Pacific Northwest of North America, it is commonly used in riparian buffer restoration and as a component of multi-story forest farming systems. In parts of Europe, particularly the UK and Scandinavia, it is integrated into hedgerows and woodland edges to support biodiversity and provide habitat. In Australia, its drought tolerance makes it suitable for integration into windbreaks and mixed farming systems in Zones 2-4, helping to protect crops and livestock from harsh conditions, and it can add biodiversity to mixed farming systems in cooler, higher rainfall regions like Tasmania or parts of Victoria. In New Zealand, it is well-suited to various temperate zones, contributing to ecological corridors and farm diversification. In Brazil, it can be planted on the edges of fields or as scattered trees to provide shade for young coffee plants and enhance biodiversity. Its resilience in cooler climates makes it a valuable asset for farmers seeking to enhance ecological function across a broad range of temperate environments.

Sources behind this view

Videos & Podcasts
Community
  • Planting blue elderberry hedgerows offers farmers dual benefits of environmental enhancement (pest control, soil health, biodiversity) and direct farm income, with potential revenue of $2,700-$4,800 p

  • A UC project is assessing the commercial viability of blue elderberries in California, exploring their potential for income generation alongside conservation benefits like hedgerows, windbreaks, and c

9

How to Integrate This Plant

Practical guidance for regenerative systems

Establishing Sambucus racemosa is typically achieved through planting nursery-grown seedlings or rooted cuttings, which is the most reliable method for ensuring genetic diversity and vigorous growth. For direct seeding, rates typically range from 1-2 lbs/acre (1.1-2.2 kg/ha) when sown in prepared beds, with a planting depth of 0.25-0.5 inches (0.6-1.3 cm). Cuttings, taken from dormant wood in late winter or early spring, can be directly planted into well-draining soil. Nursery-grown saplings, usually 1-2 feet (0.3-0.6 m) tall, are planted at a spacing of 8-15 feet (2.4-4.5 m) apart, depending on the desired density and system design. For hedgerow or windbreak plantings, spacing typically ranges from 6-10 feet (1.8-3 m) apart, allowing for mature canopy development. For alley cropping or silvopasture systems, rows can be spaced 20-40 feet (6-12 m) apart to accommodate intercropping or grazing. Planting in the Northern Hemisphere is best done in early spring (March-April) or fall (October-November), while the Southern Hemisphere follows the opposite timing (March-April or September-October).

Management practices during the establishment phase are crucial for long-term success. Young plants require consistent moisture, with approximately 1 inch (2.5 cm) of water per week during the first 1-2 years, especially during dry spells. While Sambucus racemosa is adaptable, initial fertilization can be supported by incorporating compost or well-rotted manure into the planting hole to provide a slow-release nutrient source. Fertility is best managed through biological means; incorporating compost annually, mulching with organic matter, and allowing fallen leaves to decompose will provide ample nutrients. Synthetic fertilizers should only be considered as a transitional input while biological systems are developing, and their use should be minimized over time.

Pruning in the first few years should focus on establishing a strong central leader or desired multi-stem form, removing any competing or crossing branches. This initial shaping sets the stage for future canopy management. Annual pruning in late winter or early spring, removing dead, diseased, or crossing branches, and thinning out older stems to encourage new growth, is recommended. This also helps maintain plant health, encourage fruiting, and manage light penetration for any understory plantings. A schedule of light pruning annually and heavier pruning every 3-5 years can maintain vigor. Sambucus racemosa can reach a mature height of 5-15 feet (1.5-4.5 m) and a spread of 5-10 feet (1.5-3 m).

For category-specific integration as a perennial tree or agroforestry species, Sambucus racemosa requires a thoughtful establishment and system design. Trees typically take 1-3 years to establish a robust root system and structural integrity, with significant fruit or biomass production beginning between years 3-7 and reaching full production by years 8-15. In alley cropping or silvopasture designs, rows of elderberry can be planted 30-40 ft (9-12 m) apart, allowing for equipment access and integration with grazing animals or other crops. Beneath the canopy, after the first 2-3 years of establishment, planting nitrogen-fixing ground covers like clover or vetch can enhance soil fertility and provide forage. Measurable soil carbon increases are often observed by year 5-7 as the plant matures and its root system expands. Long-term infrastructure considerations include initial irrigation for establishment and potentially deer or browse protection until the plants are well-established, especially in areas with high herbivore pressure.